State Key Laboratory of Vegetation and Environmental Change, Institute of Botany, Chinese Academy of Sciences, Beijing, China; University of Chinese Academy of Sciences, Beijing, China; Department of Ecology, College of Urban and Environmental Sciences, Key Laboratory for Earth Surface Processes, Ministry of Education, Peking University, Beijing, China; Department of Physical Geography, Stockholm University, Stockholm, Sweden; Department of Earth System Science, Stanford University, Stanford, CA, United States; Key Laboratory of Adaptation and Evolution of Plateau Biota, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining, Qinghai, China; Key Laboratory of Ecosystem Network Observation and Modeling, Institute of Geographic Sciences and Natural Resources Research, Chinese Academy of Sciences, Beijing, China; Institute of Biological and Environmental Sciences, School of Biological Sciences, University of Aberdeen, Aberdeen, United Kingdom