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population(8)
nitrogen(6)
surface water(6)
grasslands(5)
lakes(5)
vegetation(4)
biomass(3)
conservation(3)
nutrients(3)
altitude(2)
carbon(2)
density(2)
fertilizers(2)
fluorescence(2)
forests(2)
humidity(2)
oxygen(2)
phosphorus(2)
precipitation(2)
rural areas(2)
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